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I've lost track of all the details in this scenario, but some general thoughts:

If your wife pre-deceased her mother (or died within a short time of her mother (typically 30 days)), she may, under state law or the terms of MIL's Will, be ignored for the purposes of inheritance. Or MIL's Will may stipulate what happens to wife's share. If wife died sufficiently after MIL, then terms of wife's Will dictate what happens to the inheritance she is entitled to. If either person died intestate, you need to follow state law with regard to inheritance priority. Finally, any beneficiary can disclaim their inheritance, but they can't direct who will receive the disclaimed amount. If the Will doesn't address the matter, state law determines how the disclaimed amounts are distributed.

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